The Arachnid Order Solifugae

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PHYLOGENY/TAXONOMY

Phylogeny of the Solifugae
Keys to FamiliesFamily AmmotrechidaeFamily Ceromidae
Family Daesiidae
Family Eremobatidae
 

EREMOBATINAE
  Eremobates
      angustus
group
      aztecus
group
      lapazi
group
      pallipes
group
      palpisetulosus
group
       ajoanus series
        Eremobates affinis
        Eremobates ajoanus
        Eremobates bixleri
       kraepelini series
        Eremobates gracilidens
        Eremobates inkopaensis
        Eremobates kraepelini
        Eremobates otavonae
        Eremobates pallidus
        Eremobates pimanus
        Eremobates pyriflora
        Eremobates titschacki
       nodularis series
        Eremobates nodularis
       palpisetulosus series
        Eremobates bajadae
        Eremobates bantai
        Eremobates kiseri
        Eremobates marathoni
        Eremobates norrisi
        Eremobates palpisetulosus
        Eremobates polhemusi
        Eremobates texanus
       scopulatus series
        Eremobates fagei
        Eremobates girardi
        Eremobates inyoanus
        Eremobates kastoni
        Eremobates leechi
        Eremobates nanus
        Eremobates nivis
        Eremobates papillatus
        Eremobates scopulatellus
        Eremobates scopulatus
        Eremobates spissus
        Eremobates tejonus
        Eremobates tuberculatus
        Eremobates vicinus
        Eremobates villosus
        Eremobates williamsi
       unplaced
        Eremobates bajaensis

        Eremobates coahuilanus
     
  Eremobates guenini
        Eremobates hessei
        Eremobates hystrix

        Eremobates purpusi

      scaber
group
      vallis
group

  Eremocosta
  Eremorhax
  Eremothera
  Horribates

THEROBATINAE
  Chanbria
  Eremochelis
 
Hemerotrecha

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 


 

 


Family Galeodidae
Family Gylippidae
Family Hexisopodidae
Family Karschiidae
Family Melanoblossidae
Family Mummuciidae
Family RhagodidaeFamily Solpugidae
Catalog of the Solifugae

 

 
Eremobates palpisetulosus species group

Diagnosis: Muma (1951) recognized this species group as including those species of Eremobates in which the fixed cheliceral fingers of males are straight or at most slightly sinuate in dorsal view, bear a constriction but no distinct notch at their base, have an ectodorsal, toothlike process near their base, and have the mesoventral groove dilated basally. Males are also provided, he noted,  with two to 10 ctenidia on the first post-spiracular abdominal sternite. Females have roughly triangular genital opercula which are adjacent anteriorly, widely separated posteriorly, and have a deep oval pit near the ectal margin of each plate.  Muma (ibid.) reported the fondal dentition as being the same as in the scaber group, except that the females have the ectal row graded II, I, III, IV in size. He later (Muma 1986, 1987) suggested that fondal tooth formulae are not valid for genus or group separation, as they are subject to wear, especially by females.   Muma (1989), in a key to the species groups of Eremobates based on males, characterized  males of this group as having the mesal groove of the fixed cheliceral finger mesoventral in position, narrowed apically (occupying less than half of finger width), and distinctly dilated basally, with the  fixed cheliceral finger straight or at most slightly sinuate in dorsal view and with an ectodorsal, ridge-like or tooth-like process near its base. In an accompanying key to the species groups of Eremobates based on females, he characterized the females as having opercula that are widely separated posteriorly and/or slightly to distinctly lobate at their posterio-ectal angles, with a pair of deep, distinct pits about midway along their ectal margins and a posteriomesal notch that is wide and long (occupying at least a third of the opercular area) and smoothly flared or arched mesally, .

Distribution: Widely distributed in western North America. 

Included species: 

    ajoanus series
     Eremobates affinis (Kraepelin 1899)
     Eremobates ajoanus Muma & Brookhart 1988
     Eremobates bixleri Muma & Brookhart 1988
   
kraepelini series
     Eremobates gracilidens Muma 1951
     Eremobates inkopaensis  Brookhart and Cushing 2005

     Eremobates kraepelini Muma 1951
     Eremobates otavonae Muma & Brookhart 1988
     Eremobates pallidus Muma & Brookhart 1988
     Eremobates pimanus Muma & Brookhart 1988
     Eremobates pyriflora Muma & Brookhart 1988
     Eremobates titschacki (Roewer 1934)
   
nodularis series
     Eremobates nodularis Muma 1951
   
palpisetulosus series
     Eremobates bajadae Muma & Brookhart 1988
     Eremobates bantai Brookhart 1965
     Eremobates kiseri Muma & Brookhart 1988
     Eremobates marathoni Muma 1951
     Eremobates norrisi Muma & Brookhart 1988
     Eremobates palpisetulosus Fichter 1941
     Eremobates polhemusi Muma & Brookhart 1988
     Eremobates texanus Muma & Brookhart 1988
       scopulatus series
        Eremobates fagei (Roewer 1934)
        Eremobates girardi (Putnam 1883)
        Eremobates inyoanus Muma & Brookhart 1988
        Eremobates kastoni Muma & Brookhart 1988
        Eremobates leechi Muma & Brookhart 1988
        Eremobates nanus Muma 1962
        Eremobates nivis Muma & Brookhart 1988
        Eremobates papillatus Muma 1970
        Eremobates scopulatellus Muma & Brookhart 1988
        Eremobates scopulatus Muma 1951
        Eremobates spissus Muma & Brookhart 1988
        Eremobates tejonus Chamberlin 1925
        Eremobates tuberculatus (Kraepelin 1899)
        Eremobates vicinus Muma 1963
        Eremobates villosus Muma 1951
        Eremobates williamsi Muma & Brookhart 1988
      
unplaced
        Eremobates bajaensis
Muma 1986
        Eremobates coahuilanus Muma 1986
     
  Eremobates guenini (Roewer 1934)
        Eremobates hessei (Roewer 1934)
        Eremobates hystrix (Mello-Leitao 1942)

        Eremobates purpusi (Roewer 1934)

Key to subgroups:

tabular key to males of five series of species of the palpisetulosus species-group of EremobatEs banks (From muma and brookhart (1988).

Subgroups

Characters

Palpal
scopula

Fixed
finger
process
form

Fixed
finger
process
mean
percent of finger
length

Fondal notch
mean
length/width
ratio

Mean
finger
width/
fondal
notch
width
ratio

Basal
intermediate
tooth of
movable
finger

Mesal tooth
of
movable
finger

nodularis
series

absent

ridge

40.0

1.78

1.41

on primary tooth

0

palpisetulosus
series

absent

ridge

25.0 to 35.0

1.0 to 1.87

0.96 to 1.33

basal angle tooth and finger

2

ajoanus
series

absent

spur

9.0 to 15.0

0.91 to 1.51

1.08 to 1.22 

on finger margin

2

kraepelini
series

 absent

blade

20.0 to 35.0

0.73 to 1.17

1.00 to 1.46

on  finger margin

1

scopulatus
series

present

blade

15.0 to 30.0

0.73 to 1.50

1.29 to 2.50

on finger margin

0-1

Notes: Muma (1986) commented that this is proving to be the largest and most difficult species-group of Eremobates. The fondal tooth formulae, he observed, are no longer considered valid for genus or group separation, as they are subject to wear, especially by females. He further opined that females, unless collected in association with males or from within known distribution ranges, are nearly impossible to place to species.


 


 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

   
   

 



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