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EREMOBATINAE
Eremobates angustus group aztecus group lapazi group pallipes group Eremobates arizonicus Eremobates barberi Eremobates californicus Eremobates chihuaensis Eremobates cinerascens Eremobates constrictus Eremobates dentilis Eremobates dilatatus Eremobates docolora Eremobates durangonus Eremobates formicarius Eremobates gerbae Eremobates lentiginosus Eremobates pallipes Eremobates putnami Eremobates simoni Eremobates suspectus Eremobates woodruffi palpisetulosus group scaber group vallis group Eremocosta Eremorhax Eremothera Horribates THEROBATINAE Chanbria Eremochelis Hemerotrecha
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Eremobates pallipes species group
Diagnosis: Muma (1951) recognized this species group as including those species of Eremobates in which the fixed cheliceral fingers of males are straight or at most slightly sinuate in dorsal view, have a constriction but no distinct notch at the base, and lack an ectal process near the base. The mesoventral groove is distinctly dilated basally, and the first postspiracular abdominal sternite may or may not be provided with ctenidia. Females, Muma (ibid.) reported, have roughly triangular opercula that are adjacent in the anterior third to half of their length but are moderately separated posteriorly. Both rows of fondal teeth, he noted, are graded in size I, III, II, IV, although he later u(Mma 1986, 1987) suggested that fondal tooth formulae are not valid for genus or group separation, as they are subject to wear, especially by females. Brookhart and Muma (1981) characterized males of the pallipes group has having the following attributes: Fixed cheliceral finger straight or slightly curved upward with a constriction but no distinct notch or spur at the base; mesoventral groove of fixed finger dilated basally into a cup-like structure; basal flange of mesoventral groove of fixed finger distinct but variable in length, width, and pitch; movable cheliceral finger with a large principal tooth, an anterior tooth, 2 small intermediate teeth, and a mesal tooth.; palpi with or without a scopula of papillae; first post-stigmatic abdominal sternite with or without ctenidia. A/CP = 5.45-6.20. Females were characterized as having the following attributes: fixed chelicera! finger with principal and medial teeth large, an anterior tooth half as large as principal tooth, 2 intermediate teeth between principal and medial teeth, and 1 between medial and anterior teeth; movable cheliceral finger with a large principal tooth, an anterior tooth, 2 intermediate teeth, and a mesal tooth; palpi usually without a scopula and first post-stigmatic abdominal sternite without distinct ctenidia; genital opercula roughly triangular and adjacent anteriorly but slightly to moderateiy separated posteriorly; A/CP=4.55·5.50. Both sexes were characterized as having the fondal teeth graded in size I, III, II, IV. Muma (1989), in a key to the species groups of Eremobates based on males, noted that these are small species, and characterized males of this group as having the mesal groove of the fixed cheliceral finger mesoventral in position, narrow apically (occupying less than half of finger width), and distinctly dilated basally, with the fixed cheliceral finger straight or at most slightly sinuate, constricted near its base, but without an ectodorsal tooth-like process. The anterior tooth of the movable cheliceral finger, he noted, is present and normal. In an accompanying key to the species groups of Eremobates based on females, he characterized the females as having opercula that are not or are only moderately separated posteriorly, are broadly triangular in shape, without distinct pits midway along ectal margins (except for E. mimbrenus Muma), have mesal margins that are parallel anteriorly, and straight or evenly curved to the posteriomesa1 notch. which is variable in form. The posterior mesal margins, he noted, are straight or evenly curved. Distribution:
North America:
western United States
and adjacent Mexico.
Key to species: Key to the pallipes species·group of Eremobates Banks from Brookhart and Muma (1981).
Notes: When the group was erected by Muma (1951) characters utilized in distinguishing species within the group included, for males, the number of papillae in the palpal scopula, the width of the fondal notch, the number of abdominal ctenidia, and the color of the malleoli. For females, the distinguishing characthers included the form and size of the caudal notch of the genital opercula and color of the malleoli. Other characters such as the total length, the lengths of the palpi, leg I, and leg IV, general coloration patterns. curvature of the male fixed cheliceral finger, and the presence of denticules on the lower margin of the male fixed finger were also noted. Brookhart and Muma (1981) utilized, in addition to the above, length/width measurements of the male's cheliceral fondal notch; measurements of the female genital opercula; measurements of the length and width of the propeltidium; and/or the length and width of the chelicerae. Measurements were analyzed both alone and as ratios (i.e., cheliceral L/W, propeltidial L/W, and the A/CP, which was obtained by summing the length of the palpus, leg I and leg IV, and dividing that figure by the length of the propeltidium plus the length of the chelicerae). They reported that the A/CP ration was established to demonstrate the relationship of appendages (A) to body length (CP). They regarded the body length as the combined lengths of the chelicerae and the propeltidium, excluding the abdomen which is subject to size variability caused by the presence or absence of food, eggs, or semen.
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