The Arachnid Order Solifugae

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PHYLOGENY/TAXONOMY

Phylogeny of the Solifugae
Keys to FamiliesFamily Ammotrechidae
Family Ceromidae
Family Daesiidae
Family Eremobatidae
 


 

 

 

 

 

 

Family Galeodidae
Family Gylippidae
Family Hexisopodidae
Family Karschiidae
Family Melanoblossidae
Family Mummuciidae
Family RhagodidaeFamily Solpugidae
Catalog of the Solifugae

Genus Eremochelis Roewer 1934
bilobatus species group

 

 

         Muma (1951) divided his newly erected genus Therobates Muma 1951 into three species groups.  One, the bilobatus group, was recognized as consisting of a single moderate-sized species, currently recognized as Eremochelis bilobatus (Muma 1951), which was designated as the type species of Therobates.  The bilobatus group was defined as follows:

Moderate-sized species.  Males with mesal groove of fixed finger distinct, enlarged basally, and containing one or more distinct carinae or ridges.  Apical plumose bristle of flagellum complex not greatly enlarged or elongated.  Ventral tubular bristles of complex situated in a group in the fondal notch.  Females of the only species known have the genital plates lobate on the mesal margins.  Ectal row of fondal teeth graded in size I, II, III, IV; mesal row I, III, II, IV.

         Muma (1962) subsequently added those species now known as Eremochelis acrilobatus (Muma 1962) and Eremochelis plicatus (Muma 1962) to the bilobatus group.  He also added Therobates arcellus Muma 1962, males of which were later (Muma 1963) shown to be misidentifed specimens of   Eremochelis bidepressus (Muma 1951).   Females regarded as belonging to Therobates arcellus by Muma (1962) have subsequently been referred to Eremochelis insignatus Roewer 1934 (Muma 1963, 1970a).  Both Eremochelis bidepressus and Eremochelis insignatus Roewer 1934 are current members of the branchi species groupThe concept of the bilobatus group remained largely unchanged despite these additions, although Muma noted that the size gradation of the fondal teeth is variable between species and specimens, indicating instability of the character.
         Muma (1962) also recognized what he called the arcus group, characterizing it as consisting of small to moderate-sized species in which the apical plumose bristle of the flagellum complex on the chelicera of the male arches over the basal portion of the mesial groove and is neither enlarged nor flattened, the ventral tubular bristles of the complex are situated in a row just above the fondal notch, and the mesial groove of the male's fixed cheliceral finger is a narrow, shallow groove containing no distinct carinae.  The only known female was described as having opercula that are expanded and lobate at the ectocaudal angle.  Ectal and mesial rows of fondal teeth were reported to be graded I, III, II, IV or I, II, III, IV  in size.  Into the group he placed what are now known as Eremochelis arcus (Muma 1962), Eremochelis macswaini (Muma 1962), and Eremochelis cuyamacanus (Muma 1962), without altering his characterization of the group.
          Muma (1963) later affiliated the species now known as Eremochelis flexacus (Muma 1963) and Eremochelis nudus (Muma 1963) with the arcus group, along with Therobates attritus Muma 1963 (described from a female specimen), which has since (Muma 1970a) been synonymized with Eremochelis imperialis (Muma 1951), a member of the imperialis species group.    Muma (1970a) provided a tabular key to the species of the arcus group, but did not alter the diagnoses of either the arcus nor bilobatus groups.
         Muma (1986) next added Eremochelis truncus Muma 1986 and Eremochelis rossi Muma 1986 to the bilobatus group, and listed Eremochelis flexacus under the bilobatus group heading, but did not offer a revised concept of either the bilobatus or arcus groups, and noted that the newly described species "do not significantly alter" any of the species group concepts.
         Muma (1989) combined the arcus group with the bilobatus group, noting that his newly described species indicate that the groups cannot be maintained separately.  He revised his diagnosis of the bilobatus group to include taxa in which the mesoventral groove of the male's fixed cheliceral finger may or may not contain distinct carinae, and in which the female opercula may or may not be lobate mesally within the posterior notch, or have lateral pits.   In his key to species groups, Muma (ibid) characterized males of the bilobatus group as having the following characteristics:  Fixed cheliceral finger with a distinct, elongate, mesoventral or dorsomesal cavity or slot occupying at least half of finger length; apical plumose seta of flagellum complex obscurely or not widened and flattened.  Females opercula provided posteriorly with a distinct mesal notch and commonly provided with anteriolateral pits and often with adjacent distinct mesal lobes at the anterior end of the notch.  He added Eremochelis cochiseae Muma 1989, Eremochelis giboi Muma 1989, Eremochelis kerni Muma 1989, and Eremochelis noonani Muma 1989 to the newly expanded bilobatus group, and
Vázquez (1991) added Eremochelis lagunensis Vázquez 1991.

         The bilobatus species group, as currently recognized, thus consists of the following taxa:

Eremochelis acrilobatus (Muma 1962)
Eremochelis arcus
(Muma 1962)
Eremochelis bilobatus
(Muma 1951)
Eremochelis cochiseae
Muma 1989
Eremochelis cuyamacanus
(Muma 1962)
Eremochelis flexacus
(Muma 1963)
Eremochelis giboi
Muma 1989
Eremochelis kerni
Muma 1989
Eremochelis lagunensis
Vázquez 1991
Eremochelis macswaini
(Muma 1962)
Eremochelis morrisi
(Muma 1951)
Eremochelis noonani
Muma 1989
Eremochelis nudus
(Muma 1963)
Eremochelis plicatus
(Muma 1962)
Eremochelis rossi Muma 1986
Eremochelis truncus Muma, 1986

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 


 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 


 

   
   
 



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