The Arachnid Order Solifugae

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PHYLOGENY/TAXONOMY

Phylogeny of the Solifugae
Keys to FamiliesFamily Ammotrechidae
Family Ceromidae
Family Daesiidae
Family Eremobatidae
Family Galeodidae
Genus Galeodes
     Galeodes abessinicus
     Galeodes adamsi
     Galeodes afghanus
     Galeodes agilis
     Galeodes anatoliae
     Galeodes annandalei
     Galeodes arabs
        Galeodes a. arabs
        Galeodes a. syriacus
     Galeodes araneoides
     Galeodes armeniacus
     Galeodes ater
     Galeodes atriceps
     Galeodes atroluteus
     Galeodes atrospinatus
     Galeodes aulicus
     Galeodes auronitens
     Galeodes australis
     Galeodes babylonicus
     Galeodes bacillatus
     Galeodes bacillifer
     Galeodes bacilliferoides
     Galeodes bactrianus
     Galeodes barbarus
     Galeodes belutschistanus
     Galeodes bengalicus
     Galeodes bicolor
     Galeodes blanchardi
     Galeodes bocharicus
     Galeodes bogojavlenskii
     Galeodes bubi
     Galeodes caspius
        Galeodes c. caspius
        Galeodes c. fuscus
        Galeodes c. pallasi
        Galeodes c. subfuscus
     Galeodes chitralensis
     Galeodes citrinus
     Galeodes clavatus
     Galeodes conversus
     Galeodes crassichelis
     Galeodes ctenogaster
     Galeodes ctenoides
     Galeodes cursor
     Galeodes dahlahensis
     Galeodes darendensis
     Galeodes darius
     Galeodes dekanicus
     Galeodes dellacaveae
     Galeodes discolor
     Galeodes distinctus
     Galeodes edentatus
     Galeodes egregius
     Galeodes elegans
     Galeodes ephippiatus
     Galeodes excelsius
     Galeodes fatalis
     Galeodes fessanus
     Galeodes festivus
     Galeodes flavivittatus
     Galeodes forcipatus
     Galeodes franki
     Galeodes fremitans
     Galeodes fumigatus
     Galeodes graecus
     Galeodes granti
     Galeodes gravelyi
     Galeodes gromovi
     Galeodes hellenicus
     Galeodes indicus
     Galeodes inermis
     Galeodes insidiator
     Galeodes interjectus
     Galeodes intermedius
     Galeodes interritus
     Galeodes karunensis
     Galeodes kermanensis
     Galeodes koeiena
     Galeodes kozlovi
     Galeodes kraepelini
     Galeodes krausi
     Galeodes lacertosus
     Galeodes laevipalpis
     Galeodes laniator
     Galeodes lapidosus
     Galeodes lawrencei
     Galeodes levyi
     Galeodes limitatus
     Galeodes lindbergi
     Galeodes litigiosus
     Galeodes loeffleri
     Galeodes luteipalpis
     Galeodes lybicus
     Galeodes lycaonis
     Galeodes macmahoni
     Galeodes marginatus
     Galeodes mauryi
     Galeodes medusae
     Galeodes melanalis
     Galeodes melanopalpus
     Galeodes minimus
     Galeodes minitor
     Galeodes mongolicus
     Galeodes montivagans
     Galeodes mosconibronzii
     Galeodes nachitschevanicus
     Galeodes nigrichelis
     Galeodes notatus
     Galeodes olivieri
     Galeodes orientalis
     Galeodes pallescens
     Galeodes palpalis
     Galeodes parvus
     Galeodes perotis
     Galeodes philippovi
     Galeodes philippoviczi
     Galeodes pinkasi
     Galeodes pirzadanus
     Galeodes pococki
     Galeodes przevalskii
     Galeodes pugnator
     Galeodes pusillus
     Galeodes rapax
     Galeodes reimoseri
     Galeodes revestitus
     Galeodes rhamses
     Galeodes rhodicola
     Galeodes roeweri
     Galeodes rufogriseus
     Galeodes rufulus
     Galeodes ruptor
     Galeodes sabulosus
     Galeodes sarpolensis
     Galeodes schach
     Galeodes schendicus
     Galeodes scythicus
     Galeodes sedulus    
     Galeodes sejugatus
     Galeodes separandus
     Galeodes setipes
     Galeodes setulosus
     Galeodes signatus
     Galeodes simplex
     Galeodes smirnovi
     Galeodes somalicus
     Galeodes spectabilis
     Galeodes spinipalpis
     Galeodes starmuehlneri
     Galeodes striatipalpis
     Galeodes subbarbarus
     Galeodes subsimilis
     Galeodes sulfuripes
     Galeodes sulphureopilosus
     Galeodes tangkharzarensis
     Galeodes tarabulus
     Galeodes taurus
     Galeodes testaceus
     Galeodes theodori
     Galeodes tillmani
     Galeodes timbuktus
        Galeodes t. brunneipalpis
        Galeodes t. timbuktus
     Galeodes toelgi
     Galeodes trichotichnus
     Galeodes trinkleri
     Galeodes tripolitanus
     Galeodes truculentus
     Galeodes turanus
     Galeodes turcmenicus
     Galeodes turkestanus
     Galeodes turki
     Galeodes tuxeni
     Galeodes uzbecus
     Galeodes veemi
     Galeodes venator
     Galeodes ventralis
     Galeodes versicolor
     Galeodes viridipilosus
     Galeodes vittatus
     Galeodes wadaicus
     Galeodes zarudnyi 

Genus Galeodopsis
Genus Galeodumus
Genus Gluviema
Genus Othoes
Genus Paragaleodes
Genus Paragaleodiscus
Genus Roeweriscus
Genus Zombis

Family Gylippidae
Family Hexisopodidae
Family Karschiidae
Family Melanoblossidae
Family Mummuciidae
Family Rhagodidae

 Family Solpugidae
Catalog of the Solifugae

Galeodes arabs C .L. Koch 1842


 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 


 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

Galeodes arabs arabs C.L Koch

Distribution: Algeria, Djibouti, Egypt, Ethiopia, I.ran, Iraq, Israel, Kenya, Libya, Morocco, Niger, Oman, Saudi Arabia, Somalia, Sudan, Syria, Tunisia, Turkey, Yemen.

Galeodes arabs syriacus Kraepelin

Galeodes arabs syriacus Kraepelin, 1899a: 203; Kraepelin, 1901: 22 (as Galeodes arabs syriaca [sic]); Werner, 1902: 607 (as Galeodes arabs syriaca [sic]); Roewer, 1934: 532; Moritz and Fischer, 1980: 148.
Type localities: Beirut, Jabal Lubnan, Lebanon; Yafo (as Jaffa), Tel Aviv, Israel.
Distribution: Israel, Lebanon, Syria.

Type localities:
of Galeodes arabs: Arabia.
of Gluvia gracilis: Colombia (probably mislabelled) .
of Galeodes lucasii: southern Algeria (as "Algiria australiori") .

Galeodes araneoides (Pallas): Olivier, 1807: 307, plate 42 fig. 3; Blanchard, 1852: 25? (not seen) (misidentifications).
Solpuga araneoides (Pallas): Audouin, 1826: 176-178, fig. 7; Audouin, 1829: 416-419, fig. 7 (misidentifications ).
Galeodes arabs
C .L. Koch, 1842: 353; C .L. Koch, 1847: 85-86, fig. 1476; C .L. Koch, 1850: 97; L. Koch, 1878: 38; Simon, 1879a: 105; Bernard, 1894a: fig. 2; Pocock, 1895a: 77 (in part; see Galeodes graecus (C .L. Koch); Pocock, 1895b: 446-447; Bernard, 1896: 307, plate XXVII figs 5, 11, plate XXVIII fig. 15, plate XXX fig. 11, plate XXXI figs 6, 10, 16-18, plate XXXII figs 18-19, plate XXXIV fig. 10; Pavesi, 1897b: 158; Pocock, 1897a: 392; Kraepelin, 1899a: 202-203; Kraepelin, 1899b: 376; Kraepelin, 1901: 21; Pocock, 1902a: 239, fig. 27; Kraepelin, 1903: 571-572; Birula, 1905b: 253; Tullgren, 1907a: 1; Birula, 1908a: 149-150; Birula, 1909: 520-521; Simon, 1909b: 42; Hirst, 1911a: 220, fig. 1; Lampe, 1918: 207; Birula, 1925: 192-200, figs 1, 3a, 4a, Sa, 6a; Loveridge, 1925: 309; Binda, 1926: 179, 181-183; Caporiacco, 1927: 60 (in part, see Galeodila somalica Caporiacco); Borelli, 1930: 219; Caporiacco, 1932: 406; Roewer, 1933: figs 176-177, 192; Roewer, 1934: 532; Caporiacco, 1936: 98; Fage, in Fage and Simon, 1936: 309-310; Birula, 1938: fig. 74a; Warren, 1939a: 167-168, figs Ig-k, 8(6, 13-14); Warren, 1939b: fig. 9(3); Caporiacco, 1941: 42; Moriggi, 1941: 286-287; Roewer, 1941: 161; Caporiacco, 1944: 95; Zilch, 1946: 142; Millot and Vachon, 1949: figs 242, 272; Vachon, 1950: 106; Lawrence, 1954a: 120; Petrunkevitch, 1955a: fig. Ill; Kastner, 1956: fig. 588; Vachon, 1958: 477-480, figs 1-7; Benoit, 1964: 97; Levy and Shulov, 1964: 109; Savory, 1964a: figs 34, 89, 94; Cloudsley-Thompson, 1968: 108, 110, 112, 114, 115, fig. 23; Kaestner, 1968: 224, fig. 12-9; Kaestner, 1969: fig. 617; Cloudsley-Thompson, 1977: 65; Savory, 1977: figs 94, 99, 104; Moritz and Fischer, 1980: 140; Bauchhenss, 1983: 129, fig. 8; Barnes, 1987: fig. 13-44; Vanni, Bartolozzi and Whitman-Mascherini, 1988: 375; Brusca and Brusca, 1990: fig. 4h; Alroth, 1992: fig. 1; Ruppert and Barnes, 1994: fig. 13-39a; Selden and Dunlop, 1998: fig. 7.l.
Gluvia gracilis
C .L. Koch, 1842: 355; C .L. Koch 1847: 97-98, fig. 1485; C .L. Koch, 1850: 97; Butler, 1873: 424; Simon, 1879a: 144 (synonymiszed by Kraepelin, 1901: 21).
Galeodes lucasii
Dufour, 1861: 385-388, plate 2 fig. 5; Butler, 1873: 420 (synonymiszed by Simon, 1879a: 99).
Galeodes graecus
C .L. Koch: Simon, 1879a: 100 (in part).
Rhax arabs
(Olivier): Simon, 1879a: 123.
Zerbina gracilis
(C .L. Koch): Karsch, 1880: 233, figs 17, 17a-b.
Zerbina (Gluvia) gracilis
(C .L. Koch): Putnam, 1883: 268.
Galeodes (Galeodes) arabs
C .L. Koch: Birula, 1914: 662. Not Galeodes arabs C .L. Koch: Butler, 1873: 418 (misidentification, see Galeodes araneoides (Pallas)); Cloudsley-Thompson, 1961a: 145-152, figs 1-3 (misidentification; see Galeodes granti Pocock); Cloudsley-Thompson, 1961b: 257-263, figs 1-3 (misidentification; see Galeodes granti Pocock); Cloudsley-Thompson, 1964a: 453 (misidentification; see Galeodes granti Pocock). Galeodes arabs arabs C .L. Koch: Kraepelin, 1899a: 203 (as "G. arabs typicus").

Types:    .

Original description ():

Translation: .

Subsequent accounts:

Roewer 1934

Schlüssel der ♂♂ (inkl. Diagnose):
1". Beweglicher Finger mit 2 oder 3 Zwischenzähnen...................................................................8
8'. Beweglicher Finger mit 2 Zwischenzähnen..............................................................................9
9'. Unbeweglicher Finger mit 1 Zwischenzahn............................................................................10
10". Metatarsus der Pedipalpen ventral mit Zylinderborsten...........................................................l5
15"

Wenigsten das 6. Sternit de Opisthosoma mit Ctenidien besetzt............................................20

20".

Außer dem 6. Sternit des Opisthosoma auch andere Sternite mit Ctenidien besetzt................28

28".

4. Tarsus ventral mit besonders differenzierten Sohlenhaaren.................................................29

29".  

Sohlenhaare des 4 Tarsus nadelförmig (typ. I ); die Ctenidien am 5.-7. (besonders 6.) Sternit des Opisthosoma nadelförmig (typ. I); Färbung gelbbraun. Opisthosoma dorsal mit oder ohne schwarze Mittelbinde; Körperlänge 35-51 mm; Arabien. Syrien. Mesopotamien, Nordost-Afrika von Abessinien und dem östl. Sudan bis Algier..........................................49. G. arabs
 

Schlűssel der ♀♀ (inkl. Diagnose):
1". Beweglicher Finger mit 2, 3 oder 4 Zwischenzähnen.............................................................12
12'. Beweglicher Finger mit 2 Zwischenzähnen............................................................................13
13'. 

Unbeweglicher Finger mit 1 Zwischenzahn............................................................................14

14' 6. Sternite of the Opisthosoma with a single Row of needle-shaped Ctenidia (typ. I); Opisthosoma dorsally with or without dark Middle band; Body length as far as 51 mm; Arabien, Syrien, Mesopotamien,Northost-Afrika von Abessinien and dem östl. Sudan as far as Algier....................................................................................................................9. G. arabs

49. Galeodes arabs C. L. Koch 1842 S. 353; weitere Lit. und Synonyma: Kraepelin 1899 S. 202 und 1901 S. 21 (ink1. var. G. a. syriacus, nur ); Birula 1905 S. 253 (?).
, - Mesopotamien (Mosul, Amara); Syrien und Palästina (Tiberias, Sarafend bei Jaffa, Jerusalem, Aïn or Sultan); Arabien (Uwinid, Audan, Mulailih, Junaira, Dibba, Yemen); Aegypten (Alamein, Keskeh, Khartoum, Henou, Kairo, Assuan); Nil-Sudan (Mongalla, Lake Rudolph, River Kaliokwell, Fort Hall); Abessinien (Hawash Railway Obock, Massaua, Dalaimale, Segirso, Webi Mane): Kordofan; Algier, Tunis (Don, Oran, Sfax, Bizerta); Tripolis (Audjila) und Cyrenaika (Benghasi, Tripolis) - (vidi 40 , 82 , non Typus).

Translation: .

Schlüssel der ♂♂ (inkl. Diagnose):
1". Beweglicher Finger mit 2 oder 3 Zwischenzähnen..................................................................8
8'. Beweglicher Finger mit 2 Zwischenzähnen.............................................................................9
9'. Unbeweglicher Finger mit 1 Zwischenzahn...........................................................................10
10". Metatarsus der Pedipalpen ventral mit Zylinderborsten..........................................................l5
15"

Wenigsten das 6. Sternit de Opisthosoma mit Ctenidien besetzt...........................................20

20".

Außer dem 6. Sternit des Opisthosoma auch andere Sternite mit Ctenidien besetzt...............28

28".

4. Tarsus ventral mit besonders differenzierten Sohlenhaaren................................................29

29".

Hairs on the soles of the 4th tarsus needle-like (Type I ); the ctenidia on the 5th through 7th (especially 6th) sternite of the opisthosoma needle-like  (Type I); Coloration orange brown. Opisthosoma dorsally with or without black median band; Body length 35-51 mm; Arabia. Syria. Mesopotamia, Northeast-Afrika from Abyssinia and eastwards to Sudan to Algieria..............................................................................................................49. G. arabs
 

Key to females ♀♀ (including diagnoses):
1". Movable finger with 2, 3 or 4 intermediate teeth...................................................................12
12'. Movable Finger with 2 Intermediate teeth.............................................................................13
13'. 

Fixed Finger with 1 Intermediate tooth.................................................................................14

6th sternite of the opisthosoma with a single row of needle-shaped ctenidia (typ. I); Opisthosoma dorsally with or without dark median band; Body length up to 51 mm; Arabia, Syria, Mesopotamia,northeast Africa from Abyssinia and eastwards through Sudan as far as Algieria.................................................................................................................9. G. arabs
14'

Distribution: Algeria, Djibouti, Egypt, Ethiopia, Iran, Iraq, Israel, Kenya, Lebanon, Libya, Morocco, Niger, Oman, Saudi Arabia, Somalia, Sudan, Syria, Tunisia, Turkey, Yemen

 

 

 



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