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PHYLOGENY/TAXONOMY
  
 
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Eremobates mormonus Roewer 1934
Eremoperna mormona Roewer 1934:
561, figs 323e, 324f; Zilch, 1946: 149.
"Eremobates geniculatus (Simon)": Muma, 1951: 55-57, figs
54-57 (misidentification).
Eremobates mormonus (Roewer): Muma, 1963: 1; Muma, 1970a: 12, fig. 9;
Brookhart, 1972: 32; Muma, 1974a: 8; Muma, 1976: 15; Vázquez, 1981b: 80;
Punzo, 1994c: 293-297; Punzo, 1994d: 188-193; Punzo, 1995b: 109-112; ►Brookhart
and Cushing, 2004: 300-302, figs 29, 37, 51, 57; ►Brookhart and Brookhart,
2006: 314.
Eremobates mimbrenus Muma, 1989: 12-13, figs 10-13 (synonymized by ►Brookhart
and Cushing 2004: 300).
Not Eremobates mormonus (Roewer): Muma, 1951: 67-69, figs 92-94
(misidentification, see Eremobates kraepelini Muma).
HOLOTYPE:
Eremoperna mormona:
UNITED STATES: Utah.
(SMF, No.RII/3466).
Eremobates mimbrenus:
UNITED STATES:
New Mexico:
Grant County
-
Gila National Forest, Signal Peak
(32º55'N, 108º10'W),
17 June 1976 (Martin H. Muma),
♂ (holotype)
(FSCA).
Original description:
Eremoperna mormona Roewer 1934:
561, figs 323e, 324f;
SUBSEQUENT ACCOUNTS:
Brookhart and Cushing, 2004: 300-302, figs 29, 37, 51, 57:
Males: Coloration dusky straw yellow over
all, propeltidium dusky purple anteriorly and on the lateral one
third (Fig. 51), tergites with broad violet brown stripe, abdomen
grey, palpal tarsus and metatarsus dusky amber, legs dusky amber on
femur-tibia joint. FF regularly curved; MF with large PT and AT, no
cleft under AT, posterior IT in the notch of PT, MST tiny to absent.
Fondal notch wider than long (Fig. 29); ctendia 4 flat, swordlike,
extending slightly more than half the length of succeeding sternite
(Fig. 37); 80 1 papillae on
palpal scopula. Apical plumose bristle covers most of mesal groove.
Male measurements (5): Total length 15.0.–23.0, palpus length
10.0–22.0, first leg length 9.0–14.5, fourth leg length 14.5–23.0,
chelicera length 3.5–5.4, chelicera width 1.6–2.6, propeltidium
length 1.6–2.8, propeltidium width 2.00–3.85. Ratios: A/CP
6.08–7.59, CL/CW 1.94–2.20, PL/PW 0.70–0.80, FL/FW 0.50–0.67, FW/FFW
1.25–1.88, CW/FFW 4.60–6.20.
Females: Coloration same as males. Chelicera typical of species;
MF with posterior IT in the notch of PT, no cleft under AT. MST
indistinct to absent. Genital operculum similar to E. zinni
and E. socal, new species as in fig. 68, pg. 59, Muma 1951
with short arms, medial surface gently curved ending in a point as
in E. zinni and E. similis, short curved wing (Fig.
57). No papillae on metatarsus of palpus; seven tiny hairlike
ctenidia were present on one specimen. Female allotype: Total
length 19.0, chelicera length 5.1, chelicera width 2.2, propeltidium
length 2.3, propeltidium width 3.9, palpus length 12.5, first leg
length 11.0, fourth leg length 18.5. Ratios: A/CP 5.75, CL/CW
2.31, PL/PW 5.90, GOL/GOW 0.54. Female measurements (3):
Total length 18.5–21.0, chelicera length 4.40–5.02, chelicera width
2.0–2.1, propeltidium length 2.1–3.0, propeltidium width 3.4–3.6,
palpus length 11.5–12.0, first leg length 10.5–12.0, fourth leg
length 18.0–22.0. Ratios: A/CP 5.34–6.16, CL/CW 2.20–2.40,
PL/PW 0.58–0.94, GOL/GOW 0.54–0.63.
DISTRIBUTION: 'U.S.A. (Arizona, Colorado, New Mexico, Utah).
PUBLISHED
RECORds: UNITED STATES:
Arizona: Coconino
County - North rim of Grand Canyon (36º12'N,
112º03'W),
13 July 1934 (Rockefeller), 2 ♀, AMNH); 18 July 1934 (Lutz),
♂,
♀,
(AMNH); 13 June 1934 (Wilton Ivie),
♂
(AMNH); Flagstaff (35º11'N,
111º39'W),
22 July 1949 (Billy Hughes),
♀
(AMNH); Kaibob Forest (35º50'N,
112º05'W),
14 June 1934 (Ivie & Rasmussen),
♂
(AMNH). Colorado: Montezuma County, Chimney Rock (37º04'N,
108º43'W),
31 July 1973 (B. Vogel). 2
♂s,
(CU); McFee Reservoir (misspelling of McPhee Reservoir) (37º34'N,
108º34'W),
28 August 1997 (B. Jacobi), 2
♂,
2
♀
(CSU); Miller Reservoir (37º37'N,
108º27'W),
28 August 1997 (no collector data).
♂
(CSU); Mesa Verde National Monument (37º14'N,
108º28'W),
23 July 1941 (no collector data), 2 ♂,
♀
(CU).
New Mexico:
Grant County
- Gila National Forest, Signal Peak (32º55'N, 108º10'W),
17 June 1976 (Martin H. Muma), ♀ (allotype of Eremobates mimbrenus)
(FSCA).
nOTES:
Brookhart and Cushing (2004)
regarded this species as
closely related to E. similis Muma, noting that it can be
distinguished from that species by its darker color, the shape of the fondal
notch, the presence of palpal papillae, and the thickness and length of the
ctenidia. They cite personal communication from
Muma, crediting him with
stating that the 'type' of E. mormonus in the collection of MNHN is
not Koch’s type, but may be a lectotype set up by Roewer and therefore
invalid. They note that no other locality data were given but
that the species must have been collected somewhere in southwest Utah based
on the presence of other species in that area. They further noted that
they found E. mormonus in northern Arizona around the Grand Canyon,
SW Colorado and western New Mexico on mesas with sage, piñon
pine-juniper and alpine meadows, and that the Arizona specimens are darker
in overall coloration.
Brookhart and Cushing (ibid.)
report that this species was erroneously described as E.
geniculatus by Muma 1951, who later, after examining the holotype in
Paris, identified it as Roewer’s species from Utah (Eremoperna
mormona Roewer 1934).
They also note that Muma's 1989 description of the holotype of E.
mimbrenus is correct except that in his drawing of the right male
chelicera, the IT is shown on the PT yet on the type specimen itself, the IT
is separate. They note that a male and female contained in the vial
with the holotype appear to be E. mormonus, as do two males and a
female from the area housed in two other vials. Further, they
observe that Muma’s (1989) description of the female allotype of E.
mimbrenus was based on a female with three intermediate teeth on the
movable finger collected in a pitfall trap in SW New Mexico, and that a male
and a female in the same vial appear to be E. mormonus. They
therefore synonymized E. mimbrenus with
E. mormonus.
Brookhart and Brookhart (2006) included this species in an annotated
checklist of continental North American Solifugae.
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