Eremobates actenidia Muma 1989
Eremobates actenidia ►Muma 1989: 1, 7, 9, 49, figs 1-2; ►Brookhart and Cushing 2004: 303,
304, 307, 310, figs. 30, 59; ►Brookhart and
Brookhart, 2006: 311.
UNITED STATES: Utah: San Juan County
, Monument Valley,
Gouldings Trading Post (37Ί06'N,
2 June 1953
(R.E. Ryckman, R.D. Lee, C.T. Ames, C.C. Lindt, and C.T.
Christianson), 1 ♂ (holotype).
Muma 1989: 1, 7, 9, 49, figs 1-2:
MALE HOLOTYPE: Total length 18.0 mm.
CP = 6.3
deeper than wide, no
ctenidia on first post-spiracular sternite, and scopula of palpal
metatarsus composed of 70 or more papillae. Otherwise, structure
similar to that of other species of group. Cheliceral dentition as
shown in figure 2. ECCS at 140x under normal light, as shown in
figure 2; also at least 2 setae located behind and below base of
principal tooth of movable cheliceral finger.
Coloration and markings similar to those of E.
septentrionis except dusky areas are fainter and less distinct.
Only distinct markings on appendages are dusky palpal tarsi and
apical ends of palpal metatarsi, figure 1.
Fondal notch dis
Figure 1.Meso-lateral view of right male palpus;
Figure 2. Ecto-lateral view of right male chelicera. Redrawn from
Brookhart and Cushing 2004: 303, 304, 307, 310, figs. 30, 59:
Appendage and propeltidium coloration dusky yellow to brownish yellow,
propeltidium tinged brownish violet on anterior and lateral margins, palpus
tinged brownish violet on tarsus and metatarsus, legs dusky yellow.
Cheliceral FF regularly curved, MF with small triangulate AT, no cleft,
small IT with the posterior separate from the PT (Fig. 32)[sic -
should be Fig. 30], no ctenidia, 70+ palpal papillae. Male holotype:
Total length 18.0, chelicera length 4.0, chelicera width 2.3, propeltidium
length 2.3, propeltidium width 3.5, palpus length 17.0, first leg length
13.0, fourth leg length 20.0. Ratios: A/CP 7.94, CL/CW 1.74. PL/PW 0.66, FL/FW 1.02, FW/FFW 1.70,
FW/CW 6.09. Male
measurements (4): Total length 17.5021.00, chelicera length 4.926.25,
chelicera width 2.292.79, propeltidium length 2.172.83, propeltidium width
3.754.58, palpus length 16.522.0, first leg length 12.013.0, fourth leg
length 19.523.0. Ratios: A/CP 5.946.78, CL/CW 22.42, PL/PW
0.580.68, FL/FW 0.791.06, FW/FFW 1.562.00, CW/FFW 6.098.13.
Coloration as in male except the legs are lightly tinged violet
at the tibia femur joint. Chelicera typical, MF with large PT, 2 IT, large
AT, posterior IT in notch of PT, no cleft under AT, tiny to absent MST.
Genital opercula with long, thin arms, a slightly curved interior margin
ending in a lobe, wings offset, posterior margin truncate (Fig. 59).
Female measurements (5): Total length 16.020.5, chelicera length
4.85.8, chelicera width 1.82.4, propeltidium length 2.22.8, propeltidium
width 3.24.4, palpus length 14.516.0, first leg length 9.511.0, fourth
leg length 16.017.0. Ratios: A/CP 4.886.37, CL/CW 2.002.67, PL/PW 0.570.75, GOL/GOW
Fig. 30. Ectal view male right chelicerae; Fig. 59. Ventral view
female genital opercula. Scale lne = 1 mm. Redrawn from Brookhart
and Cushing 2004.
UNITED STATES: Utah -
desert grass region of San Juan County.
RECORds: UNITED STATES: Utah: San Juan
County, 6.4 Km N of Bluff (378179N,
in wet pitfall
traps, 10 June26 August 2000 (Jack & Irene Brookhart), 3 ♂,
4 ♀, (DMNS).
Muma (1989), in a key to members of the
Eremobates scaber species group, characterized this species as having
a fondal notch that is distinctly deeper than wide, dusky
apical ends on the palpal metatarsi and dusky palpal tarsi, and no ctenidia.
When discussing the A/CP
ratio, Muma notes that
the A/CP of 6.86 for male Eremobates
c1arus indicates shorter legs than Eremobates actenidia, which
has an A/CP of 7.93 for males. In diagnosing this species, Muma (ibid) reported that the lack of abdominal ctenidia
distinguishes this species from all other members of the
Eremobates scaber group, and further noted that it can be
separated from E. ascopulatus by a scopula of 70+ papillae,
that the pale legs and dusky palpal tips distinguish it from E.
septentrionis, and that it is much smaller in size than most species,
which also serves to distinguish it from
Brookhart and Cushing (2004), in a key to males of the scaber group,
characterized this species as lacking ctenidia and having a
palpal tarus and metatarsus Elsewhere,
they stated that this species may be separated from all other scaber grouip
members but E. ctenidiellus by its
lack of ctenidia on the first post stigmatal segment, noting that Eremobates
ctenidiellus generally lacks ctenidia, but its dark coloration and the shape of
the fondal notch, which is significantly longer than wide, easily
distinguish it from E. actenidia. They further observe that Eremobates
ctenidiellus also has a high
A/CP ratio indicating longer appendages and a statistically thinner male
fixed finger in relation to the fondal notch. They report that
differs significantly from others in the group and indicates a thinner fixed
cheliceral finger. In
regards to this species' distribution,
Brookhart and Cushing (2004)
actenidia has only been found in the desert grass region of San Juan
County, Utah. Brookhart collected from pitfall traps set in three different
habitats along a 16 km stretch of Hwy 195 in San Juan County, Utah from 29
May 200028 August 2000 and again on 6 June 2001. Eremobates actenidia was
collected from desert grasslands but not from desert shrub or Piρon-Juniper
assemblages in this transect,.
Eremobates mormonus was found 161 km east in Montezuma County,
Colorado and E. corpink, new species, is found 161 km west in the
Coral Pink Sand Dunes of Kane County, Utah at approximately the same
latitude. Brookhart and Brookhart (2006) include the species in
an annotated checklist of continental North American Solifugae, noting that
their were, at that time, four recorded specimens ("Four males and four